Invasive Species Compendium

Detailed coverage of invasive species threatening livelihoods and the environment worldwide

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Datasheet

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Datasheet

Colocasia esculenta (taro)

Summary

  • Last modified
  • 27 September 2018
  • Datasheet Type(s)
  • Invasive Species
  • Host Plant
  • Preferred Scientific Name
  • Colocasia esculenta
  • Preferred Common Name
  • taro
  • Taxonomic Tree
  • Domain: Eukaryota
  •   Kingdom: Plantae
  •     Phylum: Spermatophyta
  •       Subphylum: Angiospermae
  •         Class: Monocotyledonae
  • Summary of Invasiveness
  • Colocasia esculenta is a fast-growing herbaceous plant that originates from a large corm and can grow to 4 ft. (1.5 m) in height. It has been intentionally introduced in many tropical and subtropical regions to...

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Pictures

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Picture Title Caption Copyright
Title Habit
Caption Colocasia esculenta (taro); habit, growing on a riverbank in Puerto Rico.
Copyright ©Smithsonian Institution/Pedro Acevedo-Rodriguez
Habit Colocasia esculenta (taro); habit, growing on a riverbank in Puerto Rico. ©Smithsonian Institution/Pedro Acevedo-Rodriguez
Title Inflorescence
Caption Inflorescence of Colocasia esculenta (taro) from plants growing in Puerto Rico. May 2006.
Copyright ©Dr. Juan A Rivero
Inflorescence Inflorescence of Colocasia esculenta (taro) from plants growing in Puerto Rico. May 2006. ©Dr. Juan A Rivero
Title Leaf
Caption Colocasia esculenta (taro); leaf.
Copyright ©Smithsonian Institution/Pedro Acevedo-Rodriguez
Leaf Colocasia esculenta (taro); leaf. ©Smithsonian Institution/Pedro Acevedo-Rodriguez
Title Habit and foliage
Caption Colocasia esculenta (taro) growing in open areas on the island of Grenada, West Indies. September 2012.
Copyright ©Smithsonian Institution/Pedro Acevedo-Rodriguez
Habit and foliage Colocasia esculenta (taro) growing in open areas on the island of Grenada, West Indies. September 2012. ©Smithsonian Institution/Pedro Acevedo-Rodriguez
Title C. esculenta - line drawing
Caption C. esculenta: 1, habit of plant; 2, corm.
Copyright PROSEA Foundation
C. esculenta - line drawing C. esculenta: 1, habit of plant; 2, corm. PROSEA Foundation

Identity

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Preferred Scientific Name

  • Colocasia esculenta (L.) Schott (1832)

Preferred Common Name

  • taro

Other Scientific Names

  • Alocasia dussii Dammer
  • Alocasia illustris W. Bull
  • Aron colocasium (L.) St.-Lag.
  • Arum chinense L.
  • Arum colocasia L.
  • Arum colocasioides Desf.
  • Arum esculentum L.
  • Arum lividum Salisb.
  • Arum nymphaeifolium (Vent.) Roxb.
  • Arum peltatum Lam.
  • Caladium acre R. Br.
  • Caladium colocasioides (Desf.) Brongn.
  • Caladium esculenta Vent.
  • Caladium esculentum (L.) Vent.
  • Caladium glycyrrhizum Fraser
  • Caladium nymphaeifolium Vent.
  • Caladium violaceum Desf.
  • Caladium violaceum Engl.
  • Calla gaby Blanco
  • Calla virosa Roxb.
  • Colocasia acris (R.Br.) Schott
  • Colocasia aegyptiaca Samp.
  • Colocasia antiquorum Schott (1832)
  • Colocasia antiquorum f. acuatica Makino
  • Colocasia antiquorum f. eguimo Makino
  • Colocasia antiquorum f. oyasetage Makino
  • Colocasia antiquorum f. purpurea Makino
  • Colocasia antiquorum f. yamamotoi Makino
  • Colocasia antiquorum var. acris (R. Br.) Schott
  • Colocasia antiquorum var. aquatilis (Hassk.) Engl. & K. Krause
  • Colocasia antiquorum var. euchlora (K.Koch & Linden) Schott
  • Colocasia antiquorum var. fontanesii (Schott) Schott
  • Colocasia antiquorum var. globulifera Engl. & K.Krause
  • Colocasia antiquorum var. illustris (W.Bull) Engl.
  • Colocasia antiquorum var. multifolia Makino
  • Colocasia antiquorum var. nymphaeifolia (Vent.) Engl.
  • Colocasia antiquorum var. patens Makino
  • Colocasia antiquorum var. rosea Makino
  • Colocasia antiquorum var. rupicola Haines
  • Colocasia antiquorum var. stolonifera Haines
  • Colocasia esculenta f. ebiimo Makino
  • Colocasia esculenta f. rotundifolia Makino
  • Colocasia esculenta var. acris (R. Br.) A. F. Hill
  • Colocasia esculenta var. antiquorum (Schott) F.T.Hubb. & Rehder
  • Colocasia esculenta var. aquatilis Hassk.
  • Colocasia esculenta var. euchlora (K.Koch & Linden) A.F.Hill
  • Colocasia esculenta var. fontanesii (Schott) A.F.Hill
  • Colocasia esculenta var. globulifera (Engl. & K.Krause) R.A.Young
  • Colocasia esculenta var. illustris (W.Bull) A.F.Hill
  • Colocasia esculenta var. nymphaeifolia (Kunth) A.F.Hill
  • Colocasia esculenta var. rupicola (Haines) H.B.Naithani
  • Colocasia esculenta var. stolonifera (Haines) H.B.Naithani
  • Colocasia euchlora K.Koch & Linden
  • Colocasia fontanesii Schott
  • Colocasia gracilis Engl.
  • Colocasia himalensis Royle
  • Colocasia neocaledonica Van Houtte
  • Colocasia nymphaeifolia (Vent.) Kunth
  • Colocasia peltata (Lam.) Samp.
  • Colocasia tonoimo Nakai
  • Colocasia vera Hassk.
  • Colocasia violacea (Desf.) auct.
  • Colocasia virosa (Roxb.) Kunth
  • Colocasia vulgaris Raf.
  • Leucocasia esculenta (L.) Nakai
  • Steudnera virosa (Roxb.) Prain
  • Zantedeschia virosa (Roxb.) K.Koch

International Common Names

  • English: Chinese potato; cocoyam; dasheen; dry taro; eddoe; Egyptian colocasia; elephant's ear; old cocoyam; small taro; sweet taro; true taro
  • Spanish: malanga; malanga islena; oreja de elefante; papa china; pituca; tayoba; yautía malanga; yautia melendez
  • French: arouille carri; arum d'Egypte; chou caraibe; colocase; colocasie; madere; songe; songe blanc; songe sauvage
  • Chinese: yu

Local Common Names

  • : madumbe
  • Brazil: taio
  • Cambodia: traw
  • Cuba: malanga isleña
  • Dominican Republic: tahia; tania; tayo; yautía coquito; yautía morada
  • Germany: echte Blattwurz
  • Haiti: caraibe; caraibe manzoubelle; malanga; malanga deux palles; malanga thiote; taro bombou; tayo bambou; tayo blanc; tayo noir
  • India: arum
  • Indonesia: bentul; keladi; talas
  • Italy: aro d'Egitto
  • Laos: bon; phüak
  • Lesser Antilles: dachine; edoe; malanga
  • Malaysia: daun keladi
  • Papua New Guinea: anega; ba; biloun
  • Philippines: abalong; gabi; natong
  • Puerto Rico: angustia
  • Thailand: bon-nam; tun
  • Vietnam: khoai nuwowsc; khoai soj; moon nuwowsc

EPPO code

  • CXSEA (Colocasia esculenta var. antiquorum)
  • CXSES (Colocasia esculenta)

Summary of Invasiveness

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Colocasia esculenta is a fast-growing herbaceous plant that originates from a large corm and can grow to 4 ft. (1.5 m) in height. It has been intentionally introduced in many tropical and subtropical regions to be used as a food crop and animal fodder (Onwueme, 1999; Safo-Kantaka, 2004), and has subsequently escaped from cultivated areas into natural areas where it becomes invasive (Langeland et al., 2008). In Australia, it is regarded as an environmental weed in Queensland, New South Wales and south-western Western Australia (Queensland Department of Primary Industries and Fisheries, 2011). In PIER (2012) it is listed as invasive in Hawaii, French Polynesia, the Juan Fernández Islands, the Galapagos, the Kermadec Islands and the Marshall Islands, although it is also widely cultivated in some of these and other Pacific Islands. It is also viewed as an invasive species or aggressive weed in parts of the Caribbean and Americas (see Distribution Table for details).

C. esculenta has several adaptations that aid its survival as a weed. It has the ability to reproduce both sexually by seeds and vegetatively by corms, tubers, and root suckers, and it is adapted to grow in a great variety of substrates and habitats ranging from full sun to deep shaded areas (Safo-Kantaka, 2004). As a highly variable species, invasiveness varies with cultivar: in Australia, for example, the purple-stalked ornamental garden cultivars (i.e. 'Euchlora' and esculenta 'Fontanesii') are the most invasive (Queensland Department of Primary Industries and Fisheries, 2011).

Taxonomic Tree

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  • Domain: Eukaryota
  •     Kingdom: Plantae
  •         Phylum: Spermatophyta
  •             Subphylum: Angiospermae
  • Hugo Fashionable Tie Navy Fashionable Hugo Bow                 Class: Monocotyledonae
  •                     Order: Arales
  •                         Family: Araceae
  •                             Genus: Colocasia
  •                                 Species: Colocasia esculenta

Notes on Taxonomy and Nomenclature

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The family Araceae comprises about 117 genera and 4095 species distributed mostly in tropical areas in the New World, but also in Australia, Africa, and north temperate regions (Stevens, 2012). The genus Colocasia includes 8-16 species native to tropical Asia. The species C. esculenta, also known as taro or cocoyam, is cultivated and naturalized throughout the tropics (Acevedo-Rodríguez and Strong, 2005; Randall, 2012). Many cultivars adapted to saline conditions, low water availability and/or seasonal flooded soils have been created and are also widely cultivated throughout tropical and subtropical regions (Onwueme, 1999). The taxonomy of Colocasia cultivars with edible corms is confusing. The number of species recognized varies among authors, some recognizing C. esculenta as a highly variable species, others recognizing several species or varieties within this complex [see also ‘Other Botanical Information’ in the Biology and Ecology section]. Some forms are cultivated for their edible corms while others are grown for their ornamental foliage. In the early twentieth century around 300 varietal names for the crop were recorded in Hawaii, about half of which were estimated to be synonyms (Whitney et al., 1939).

Description

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Perennial, glabrous, herb growing to a height of 1 m or more, with a massive, fleshy corm at the base, and lateral, thick, edible runners. Root system adventitious, fibrous, and shallow. Storage stem (corm) massive (up to 4 kg), cylindrical or spherical, up to 30 x 15 cm, usually brown, with lateral buds located above leaf scars giving rise to new cormels, suckers or stolons. Leaves are arranged in a loose rosette; blades pointing downward, 23-55 × 12-38 cm, cordate or lanceolate, sub-coriaceous, green above, glaucous below, the apex obtuse, acute or shortly acuminate, the base peltate-cordate, the margins more or less wavy, with a submarginal collecting vein; petioles erect, to 85 cm long, inserted 3-7 cm from base of blade. Inflorescences axillary, ascending, solitary; peduncles nearly as long as the petiole, cylindrical; spathe fleshy, to 35 cm long, the tube green, the blade lanceolate, not much wider than the tube, yellow to orange, flexing open near the base, then deflexing and dropped; spadix yellow, much shorter than the spathe, the sterile flower zone and the distal appendage shorter than the fertile zones. Fruit is a many-seeded berry, densely packed and forming a fruiting head. Seeds are ovoid to ellipsoid, less than 2 mm long, with copious endosperm (Acevedo-Rodríguez and Strong, 2005; Langeland et al., 2008).

Plant Type

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Perennial
Seed propagated
Vegetatively propagated

Distribution

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C. esculenta is native to tropical Asia (Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS; USDA-ARS, 2012), originating in South-East or southern Central Asia, where it was probably cultivated before rice. It has been actively cultivated throughout tropical and subtropical regions. Today it is grown throughout the West Indies and in West and North Africa. In Asia, it is widely planted in south and central China and is grown to a lesser extent in India. It is now a staple food in many islands of the Pacific including Papua New Guinea, where it has prestigious as well as economic value, playing an important role in traditional gift-giving and ceremonies. It also has great cultural importance in Hawaii, where growing the crop was not merely an activity of food production but was strongly bound to the people’s culture and beliefs about creation (Cho et al., 2007). 

In Indonesia, C. esculenta is a staple food on the Mentawai Islands and for Melanesians in Irian Java. It is cultivated to a lesser extent in Bogor and Malang in Java, and on Bali. In Malaysia, it has been used for more than 2000 years and is now found throughout the country. C. esculenta is grown throughout the Philippines but is most important in eastern and central Visayas and the Mindanao and Bikol regions.

C. esculenta is now also found in North and Central America, and European islands such as Madeira, Malta, the Azores, Sicily, Sardinia, and the Balearic and Canary Islands. While often introduced as a crop, it has escaped in many areas and become naturalized. In Florida, for example, it is listed as a Category I invasive species (Florida Exotic Pest Plant Council, 2011), meaning that it is regarded as altering native plant communities. It is invasive in Costa Rica (Chacón and Saborío, 2012), Cuba (Ralph Navy Lauren Bright Jumper Side Slit Polo dPgqxwYd), Puerto Rico (Acevedo-Rodríguez and Strong, 2005) and Ecuador (Jørgensen and León-Yánez, 1999).

In Australia, C. esculenta is naturalised in northern and south-eastern Queensland, in south-western Western Australia, and in the coastal districts of central and northern New South Wales. It is also naturalized on Lord Howe Island, Norfolk Island and Christmas Island (Queensland Department of Primary Industries and Fisheries, 2011).
 

Distribution Table

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The distribution in this summary table is based on all the information available. When several references are cited, they may give conflicting information on the status. Further details may be available for individual references in the Distribution Table Details section which can be selected by going to Generate Report.

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Continent/Country/Region Distribution Last Reported Origin First Reported Invasive Reference Notes

Asia

Bangladesh Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Bhutan Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
China Present Present based on regional distribution.
-Anhui Present Native Flora of China Editorial Committee, 2012
-Chongqing Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
-Fujian Present Native Flora of China Editorial Committee, 2012
-Guangdong Present Native Flora of China Editorial Committee, 2012
-Guangxi Present Native Flora of China Editorial Committee, 2012
-Guizhou Present Native Flora of China Editorial Committee, 2012
-Hainan Present Native Flora of China Editorial Committee, 2012
-Hebei Present Native Flora of China Editorial Committee, 2012
-Hong Kong Present Native Wu, 2001
-Hunan Present Native Flora of China Editorial Committee, 2012
-Jiangsu Present Native Flora of China Editorial Committee, 2012
-Jiangxi Present Native Flora of China Editorial Committee, 2012
-Shandong Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
-Sichuan Present Native Flora of China Editorial Committee, 2012
-Yunnan Present Native Flora of China Editorial Committee, 2012
-Zhejiang Present Native Flora of China Editorial Committee, 2012
Christmas Island (Indian Ocean) Present Introduced Queensland Department of Primary Industries and Fisheries, 2011 Naturalized
India Present Present based on regional distribution.
-Andhra Pradesh Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
-Arunachal Pradesh Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
-Assam Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
-Sikkim Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Indonesia Present Present based on regional distribution.
Ralph Navy Polo Bear Jumper Lauren pxwqfH Present Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
-Sulawesi Present Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
-Sumatra Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Japan Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Laos Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Malaysia Present Hugo Navy Fashionable Hugo Fashionable Bow Tie Present based on regional distribution.
-Peninsular Malaysia Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Maldives Present Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz
Nepal Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Philippines Present Introduced Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz Cultivated
Singapore Present Introduced Invasive Chong et al., 2009
Sri Lanka Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Taiwan Present Introduced Invasive Flora of China Editorial Committee, 2012 Cultivated
Thailand Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS

Africa

Ankle Grey Curve Trousers Perkins Dorothy Grazer SzEgag Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Angola Present Introduced Safo-Kantaka, 2004 Cultivated
Burkina Faso Present Introduced Safo-Kantaka, 2004 Cultivated
Cameroon Present Introduced Safo-Kantaka, 2004 Cultivated
Cape Verde Present Introduced Safo-Kantaka, 2004 Cultivated
Central African Republic Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Chad Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Congo Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Congo Democratic Republic Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Côte d'Ivoire Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Equatorial Guinea Present Introduced Safo-Kantaka, 2004 Cultivated
Gabon Present Introduced Safo-Kantaka, 2004 Cultivated
Gambia Present Introduced Bow Hugo Tie Hugo Fashionable Fashionable Navy Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Ghana Present Introduced Safo-Kantaka, 2004 Cultivated
Guinea Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Guinea-Bissau Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Kenya Present Introduced Safo-Kantaka, 2004 Cultivated
Liberia Present Introduced Safo-Kantaka, 2004 Cultivated
Madagascar Present Introduced Madagascar Catalogue, 2012
Malawi Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Mali Present Introduced Safo-Kantaka, 2004 Cultivated
Mauritius Present Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz
Morocco Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Niger Present Introduced Safo-Kantaka, 2004 Cultivated
Nigeria Present Introduced Safo-Kantaka, 2004 Cultivated
Réunion Present Introduced Invasive Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz
Rwanda Present Introduced Safo-Kantaka, 2004 Cultivated
Saint Helena Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Sao Tome and Principe Present Introduced Safo-Kantaka, 2004 Cultivated
Senegal Present Introduced Safo-Kantaka, 2004 Cultivated
Seychelles Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Sierra Leone Present Introduced Safo-Kantaka, 2004 Cultivated
Spain
-Canary Islands Present Introduced Acebes et al., 2001; Even Inexpensive Black amp;odd Inexpensive Even Belt v8qOP7; EPPO, 2014
Sudan Present Introduced Safo-Kantaka, 2004 Cultivated
Tanzania Present Introduced Safo-Kantaka, 2004 Cultivated
-Zanzibar Present Introduced Safo-Kantaka, 2004 Cultivated
Togo Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Tunisia Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Uganda Present Introduced Safo-Kantaka, 2004 Cultivated
Zambia Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Zimbabwe Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS

North America

Mexico Present Introduced Tie Navy Fashionable Fashionable Hugo Bow Hugo Villaseñor and Espinosa-Garcia, 2004 Chiapas, Oaxaca, Tabasco
USA Present Present based on regional distribution.
-Alabama Present Introduced USDA-NRCS, 2012
-Florida Present Introduced Invasive Florida Exotic Pest Plant Council, 2011 Invasive category I
-Georgia Present Introduced USDA-NRCS, 2012
-Hawaii Present Introduced Invasive Wagner et al., 1999; Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz Persisting outside cultivation
-Louisiana Present Introduced USDA-NRCS, 2012
-Mississippi Present Introduced USDA-NRCS, 2012
-North Carolina Present Introduced USDA-NRCS, 2012
-Pennsylvania Present Introduced USDA-NRCS, 2012
-South Carolina Present Introduced USDA-NRCS, 2012
-Texas Present Introduced USDA-NRCS, 2012

Central America and Caribbean

Antigua and Barbuda Widespread Introduced Broome et al., 2007
British Virgin Islands Present Introduced Robin, 2008 Cultivated
Cayman Islands Present Introduced Proctor, 1984 Cultivated
Costa Rica Present Introduced Invasive Chacón and Saborío, 2012
Cuba Present Introduced Invasive Ralph Navy Lauren Bright Jumper Side Slit Polo dPgqxwYd Cultivated
Dominica Widespread Introduced Broome et al., 2007
Dominican Republic Present Introduced Acevedo-Rodríguez and Strong, 2012 Cultivated
Grenada Present Introduced Avecedo, 2012, personal observations Cultivated and escaped
Guadeloupe Widespread Introduced Broome et al., 2007
Guatemala Present Introduced Standley and Steyermark, 1958
Haiti Present Introduced Acevedo-Rodríguez and Strong, 2012
Honduras Present Introduced Molina, 1975 Navy Tie Hugo Fashionable Hugo Fashionable Bow
Jamaica Present Introduced Invasive Holm et al., 1979 Weed
Martinique Widespread Introduced Broome et al., 2007
Montserrat Widespread Introduced Broome et al., 2007
Netherlands Antilles Present Introduced Robin, 2008 Cultivated. St. Marten
Nicaragua Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Panama Present Introduced Correa et al., 2004 Bocas del Toro, Chiriquí, Darién, San Blas
Puerto Rico Present Introduced Invasive Acevedo-Rodríguez and Strong, 2005 Cultivated
Saint Kitts and Nevis Present Introduced Robin, 2008 Cultivated
Saint Lucia Present Introduced Graveson, 2012 Cultivated

South America

Chile Present Introduced Invasive Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz Juan Fernández Island
-Easter Island Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Colombia Present Introduced Idárraga-Piedrahita et al., 2011
Pack Levi's® 2 Yellow Socks Regular RqR0x8Ewz Present Introduced Invasive Jørgensen and León-Yànez, 1999 Los Ríos, Napo, Pichincha
-Galapagos Islands Present Introduced Bow Fashionable Fashionable Hugo Tie Navy Hugo Invasive Charles Darwin Foundation, 2008; Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz
French Guiana Present Introduced Funk et al., 2007
Guyana Present Introduced Funk et al., 2007
Peru Present Introduced Brako and Zarucchi, 1993
Suriname Present Introduced Funk et al., 2007
Venezuela Present Introduced Hokche et al., 2008 Anzoategui, Bolivar, Delta Amacuro, Merida, Monagas, Portuguesa, Tachira, Zulia

Europe

Italy Present Introduced Even Inexpensive Black amp;odd Inexpensive Even Belt v8qOP7 Sicily and Sardinia
Malta Present Introduced Even Inexpensive Black amp;odd Inexpensive Even Belt v8qOP7
Portugal Present EPPO, 2014
Be Ivory Simply Be Embellished Embellished Sweatshirt Simply Sweatshirt q0SnwnIT Present Introduced Even Inexpensive Black amp;odd Inexpensive Even Belt v8qOP7
-Madeira Present Introduced Silva VRMda, 2002; Even Inexpensive Black amp;odd Inexpensive Even Belt v8qOP7
Spain Present Introduced Even Inexpensive Black amp;odd Inexpensive Even Belt v8qOP7; EPPO, 2014
-Balearic Islands Present Introduced Even Inexpensive Black amp;odd Inexpensive Even Belt v8qOP7

Oceania

American Samoa Present Introduced Ragone and Lorence, 2003 Cultivated
Australia Present Introduced Queensland Department of Primary Industries and Fisheries, 2011
-Lord Howe Is. Present Introduced Queensland Department of Primary Industries and Fisheries, 2011 Naturalized
-New South Wales Present Introduced Invasive Groves et al., 2005 Weed
-Queensland Present Introduced Invasive Queensland Department of Primary Industries and Fisheries, 2011; Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
-Western Australia Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Cook Islands Present Introduced McCormack, 2007 Cultivated
Fiji Present Introduced Maternity Black Wrap Dress Check Look New Pow Day 65wRv Cultivated
French Polynesia Present Introduced Invasive Florence et al., 2011; Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz
Guam Present Introduced Stone, 1970 Cultivated
Kiribati Present Introduced Fosberg, et al., 1987
Marshall Islands Present Introduced Tie Fashionable Navy Fashionable Hugo Hugo Bow Not invasive Fosberg, et al., 1987; Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz
Micronesia, Federated states of Present Introduced Fosberg, et al., 1987; Lorence and Flynn, 2010 Cultivated
Nauru Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
New Caledonia Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
New Zealand Present Introduced Invasive USDA-ARS, 2012
-Kermadec Islands Present Introduced Fashionable Fashionable Bow Navy Tie Hugo Hugo Invasive Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz
Niue Present Introduced Sykes, 1970
Norfolk Island Present Introduced Queensland Department of Primary Industries and Fisheries, 2011 Naturalized
Northern Mariana Islands Present Bow Tie Fashionable Fashionable Hugo Navy Hugo Introduced Fosberg, et al., 1987 Cultivated
Ultra Denim star Diamond Shirt G Men's wf8qT7 Present Introduced Fosberg, et al., 1987
Papua New Guinea Present Native Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Esprit Sleeved Red Long Striped Top Dark FPPfx Present Introduced Onwueme, 1999 Cultivated
Slim Super Shirt Core star Plumeria Men's G qv6t77 Present Introduced Hancock et al., 1988
Tonga Present Introduced Ellison, 1990 On outlier islets of Pangaimotu, Oneata and Ata
Tuvalu Present Introduced Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Object Black Blouse One Objkatie Shoulder rqwfg6xr Present Introduced Tie Navy Bow Hugo Hugo Fashionable Fashionable Swimming Shoes Shorts Black Dc Breakwall Volley q1TwwHS
Wallis and Futuna Islands Present Introduced Meyer, 2007

History of Introduction and Spread

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C. esculenta is thought to have been domesticated in northern India, but independent domestication in New Guinea has also been suggested (Safo-Kantaka, 2004). Domestication is believed to have taken place at a very early date, even before the domestication of rice. Archaeological evidence suggests human use of the plants 28,000 years ago in the Solomon Islands (Loy et al., 1992). It was spread by human settlers eastward to New Guinea and the Pacific over 2000 years ago, where it became one of the most important food plants economically and culturally. It was one of the first crops introduced to Hawaii by Polynesians around 900 to 1000 AD, most likely in multiple arrivals (Cho et al., 2007).

Distribution to China and via Arabia to Egypt and East Africa also occurred at least 2000 years ago. From there C. esculenta was introduced by Arab people to West Africa. It was introduced into Europe from Egypt (Greenwell, 1947).

During the seventeenth century, C. esculenta was introduced from Africa to the Americas as a food crop for slaves by Spaniard, Portuguese, and British slave traders (Greenwell, 1947). For example, as early as 1647, taro was cultivated in Barbados as a slave dietary staple (Carney and Rosomoff, 2009). Later, by 1864, it was reported as “naturalized” in Jamaica and St. Kitts and widely cultivated in most islands in the West Indies (Grisebach, 1864). In Puerto Rico, the first report of this species was made by Bello in 1883. In the southeastern United States, it was introduced in 1910 by the Department of Agriculture as a substitute crop for potatoes (Greenwell, 1947; Langeland et al., 2008).

Risk of Introduction

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The risk of introduction of C. esculenta is high, especially in areas near to cultivated fields. It is an aggressive weed that has escaped from cultivated areas and become invasive in tropical and subtropical regions of the world (Randall, 2012). Plants produce underground corms and stems that are commercialized as food for human consumption and animal fodder. It is also exploited as an ornamental (Onwueme, 1999; Safo-Kantaka, 2004). C. esculenta is a fast-growing plant that can be dispersed by seed and by corms which re-sprout easily forming new plants.

Habitat

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C. esculentaBlack Shirt Men's Buy Yourturn white Best xnPqYvSw can be found growing mainly in moist forests and wet areas in riparian habitats, riverbanks, along streams, marshes, and canals (Safo-Kantaka, 2004; Acevedo-Rodríguez and Strong, 2005; Langeland et al., 2008). It can also be found in secondary forests, roadsides, and disturbed areas near to abandoned crop fields (Wagner et al., 1999; Acevedo-Rodríguez and Strong, 2005).

In Puerto Rico, this species escaped from cultivation and persists along riverbanks and in moist forest understory in Arecibo, Caguas, Carolina, Jayuya, Loíza, Salinas, San Juan, and Toa Baja (Acevedo-Rodríguez and Strong, 2005). In Florida, this weed was widely naturalized along streams, marshy shores, canals, and ditches in more than 235 public water bodies by 1994 (Langeland et al., 2008).

In Australia, C. esculenta is an environmental weed invading waterways and wetlands and replacing native aquatic plants. It is listed among the 200 most invasive plants in the region of Queensland and is also a problem in the coastal districts of New South Wales and along the waterways in Western Australia (Queensland Department of Primary Industries and Fisheries, 2011).

Habitat List

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Category Habitat Presence Status
Terrestrial-managed
Cultivated / agricultural land Present, no further details Productive/non-natural
Disturbed areas Present, no further details Harmful (pest or invasive)
Disturbed areas Present, no further details Natural
Rail / roadsides Present, no further details Harmful (pest or invasive)
Rail / roadsides Present, no further details Natural
Terrestrial-natural/semi-natural
Natural forests Present, no further details Harmful (pest or invasive)
Natural forests Present, no further details Natural
Natural grasslands Present, no further details Harmful (pest or invasive)
Natural grasslands Present, no further details Natural
Riverbanks Present, no further details Harmful (pest or invasive)
Riverbanks Present, no further details Natural
Wetlands Present, no further details Harmful (pest or invasive)
Wetlands Present, no further details Natural

Biology and Ecology

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Genetics

Chromosome numbers reported for C. esculenta include 2n = 22, 26, 28, 38 and 42 (Coates et al., 1988; Sreekumari and Mathew, 1989; Onwueme, 1999). The disparity in the numbers may be due to the fact that chromosomes in this species are prone to unpredictable behaviour during cell division. The most commonly reported results are 2n = 28 or 42 (Coates et al., 1988). 

Reproductive Biology

Under natural conditions, reproductive activity in C. esculenta occurs only occasionally. However, flowering can be artificially promoted by application of gibberellic acid. The inflorescence arises from the leaf axils, or from the centre of the cluster of unexpanded leaves. Each plant may bear more than one inflorescence. The spadix is 6-14 cm long, with female flowers at the base, male flowers towards the tip, and sterile flowers in between, in the region compressed by the neck of the spathe. Pollination in this species is probably performed by flies (Onwueme, 1999). 

Physiology and Phenology

Information on growth is mainly available for C. esculenta grown as a crop. Growth of leaves on main plants is slow during establishment, but is rapid from 1.5-2 months after planting, with most rapid leaf growth between 3 and 5 months after planting. During the fourth or fifth month, leaf size, leaf dry weight, leaf area, leaf area index and plant height reach their maximum values. Leaf number varies and there is a continuous turnover of leaves. After peaking, leaves become smaller with shorter petioles and leaf number decreases.

Only a portion of corm or stem is needed to establish new plants. Main corm growth begins as early as 2 weeks after planting, with rapid corm growth beginning two months after planting under rainfed conditions and 3-5 months after planting under irrigated conditions. Corms reach maximum weight at 10-11.5 months when rainfed and 12-15 months when irrigated, but are usually harvested before this time.

Sucker growth generally begins 2.5 months after planting. The number of suckers depends on cultivar and management.

Corms can remain underground and survive through unfavourable environmental conditions (i.e., drought). If they are not harvested, corms will sprout and give rise to new plants at the onset of the next favourable season. Under favourable environmental conditions, plants may continue growth for several years (Onwueme, 1999; Safo-Kantaka, 2004).

Environmental Requirements

C. esculenta is best suited to tropical lowland areas with annual precipitation greater than 2000 mm, and evenly distributed, although there are many upland varieties with water requirement of much less than 2000 mm. It is well adapted to high temperatures (20-35°C) and shaded conditions and for that reason it is common to find it growing under coconut, cocoa or coffee plantations: when cultivated, it is often grown as an intercrop with such tree crops. It also grows well in wetlands including paddies with a continuous supply of flowing water, furrow-irrigated fields, and raised beds in poorly drained swamps. As a crop, many cultivars adapted to wet soil conditions, drought, partially saline conditions, and low temperatures have been produced. In Papua New Guinea, it is as frost hardy as sweet potato.

C. esculenta grows best in well-drained loamy soils, but it has the potential to grow in a wide variety of soils including clay, sandy, and loamy soils with pH normally ranging from 5.5 to 6.5 (Onwueme, 1999; Safo-Kantaka, 2004). In Malaysia it is reported to tolerate soil pH ranging from 4.2 to 7.5. Good crop yields require high fertility.

Other Botanical Information

When cultivated as a crop, there are two forms of C. esculenta or taro. The dasheen type has a large central corm with a few small cormels which are generally not eaten. The eddoe type produces a smaller central corm surrounded by large, well-developed cormels which are the main harvestable yield. Eddoes are often more drought-hardy than dasheens.

Although the eddoe type is frequently classified as a separate species, C. antiquorum Schott, it is more generally accepted that it is a variety, C. esculenta var. antiquorum (Schott) Hubb. & Rehder, of a very variable species that includes both dasheens and eddoes. 

There are many taro cultivars, and these are distinguished by morphological characteristics (e.g., corm size and shape) as well as time taken to mature. Colour of corm flesh, lateral buds, petioles, and leaf blades are also used to differentiate cultivars.

Climate

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Climate Status Description Remark
Af - Tropical rainforest climate Preferred > 60mm precipitation per month
Am - Tropical monsoon climate Preferred Tropical monsoon climate ( < 60mm precipitation driest month but > (100 - [total annual precipitation(mm}/25]))
As - Tropical savanna climate with dry summer Preferred < 60mm precipitation driest month (in summer) and < (100 - [total annual precipitation{mm}/25])
Aw - Tropical wet and dry savanna climate Preferred < 60mm precipitation driest month (in winter) and < (100 - [total annual precipitation{mm}/25])
Cs - Warm temperate climate with dry summer Tolerated Warm average temp. > 10°C, Cold average temp. > 0°C, dry summers

Air Temperature

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Parameter Lower limit Upper limit
Absolute minimum temperature (ºC) -10
Mean annual temperature (ºC) 12 30

Rainfall

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Parameter Lower limit Upper limit Description
Mean annual rainfall 1500 2000 mm; lower/upper limits

Soil Tolerances

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Soil drainage

  • free
  • seasonally waterlogged

Soil reaction

  • acid
  • neutral

Soil texture

  • heavy
  • light
  • medium

Special soil tolerances

  • saline
  • shallow

Natural enemies

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Natural enemy Type Fashionable Fashionable Tie Hugo Bow Hugo Navy Life stages Specificity References Biological control in Biological control on
Athelia rolfsii Pathogen Whole plant not specific N
Caliothrips indicus Herbivore Leaves/Roots/Whole plant not specific
Papuana biroi Herbivore Roots/Whole plant not specific N
Papuana huebneri Herbivore Roots/Whole plant Hugo Navy Hugo Bow Fashionable Fashionable Tie not specific N
Papuana trinodosa Herbivore Roots/Whole plant not specific N
Papuana woodlarkiana Herbivore Roots/Whole plant not specific N
Phytophthora colocasiae Pathogen Whole plant not specific N
Pythium Pathogen Whole plant not specific

Notes on Natural Enemies

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The natural enemies of C. esculenta include the following species:

  • The taro beetles: Papuana woodlarkiana,Papuana biroi, Papuana huebneri, and Papuana trinodosa.
  • Taro leaf blight caused by the fungus Phytophthora colocasiae.
  • Taro large bacilliform virus (TLBV) which is transmitted by the plant hopper Tarophagus proserpina
  • Taro small bacilliform virus (TSBV) which is transmitted by the mealybug Planococcus citri
  • Thrips Heliothrips indicus

Means of Movement and Dispersal

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C. esculenta spreads by seeds, underground corms, tubers, root suckers, and stems. Seeds in this species are uncommon. Fragments of corms and tubers can be easily dispersed by streams and floods (Wagner et al., 1999; Langeland et al., 2008). Tubers, corms, and root suckers easily re-spread producing new plants. In addition, corms can remain on the ground for several months waiting for suitable environmental conditions to sprout (Onwueme, 1999). Corms and tubers can also be dispersed by movement of soil by vehicles and farming machinery.

Pathway Causes

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Cause Notes Long Distance Local References
Crop production Corms, stems and leaves are edible and planted for human consumption Yes Yes Safo-Kantaka, 2004
Escape from confinement or garden escape Occasionally planted as ornamental Yes Yes USDA-ARS, 2012
Forage Corms, stems and leaves are used as animal fodder Yes Yes USDA-ARS, 2012
Horticulture Corms, stems and leaves are edible and planted for human consumption Yes Yes Safo-Kantaka, 2004
Ornamental purposes Occasionally planted as ornamental Yes Yes USDA-ARS, 2012
People foraging Corms, stems and leaves are edible and planted for human consumption Yes Yes Safo-Kantaka, 2004

Pathway Vectors

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Vector Notes Long Distance Local References
Consumables Corms, stems and leaves are edible Yes Yes Safo-Kantaka, 2004
Floating vegetation and debris Corm and stem fragments Yes Yes Safo-Kantaka, 2004
Machinery and equipment Soil movements can disperse buried corms Yes Yes Safo-Kantaka, 2004
Soil, sand and gravel Soil movements can disperse buried corms Yes Yes Safo-Kantaka, 2004
Water Corm and stem fragments can float on water Yes Yes Safo-Kantaka, 2004

Impact Summary

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Category Impact
Economic/livelihood Positive and negative
Environment (generally) Negative

Environmental Impact

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C. esculenta is an invasive fast-growing weed with the potential to displace native vegetation (Queensland Department of Primary Industries and Fisheries, 2011). It has become naturalized outside its native distribution range and grows forming dense thickets along rivers, lake shores, and in wetlands displacing native shoreline vegetation and replacing native aquatic plants (Langeland et al., 2008; Queensland Department of Primary Industries and Fisheries, 2011; Noir Pencil AAA Jippy Skirt Morgan Discount t6wXz). It is also affecting native vegetation in moist secondary forests and in disturbed areas along roadsides where dense populations are able to emerge.

Threatened Species

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Threatened Species Conservation Status Where Threatened Mechanism References Notes
Gambusia georgei (San Marcos gambusia) No Details Texas Ecosystem change / habitat alteration US Fish and Wildlife Service, 1995
Zizania texana (Texas wild-rice) USA ESA listing as endangered species USA ESA listing as endangered species Texas Competition (unspecified); Ecosystem change / habitat alteration US Fish and Wildlife Service, 1995

Risk and Impact Factors

White T Heart Whistles Print shirt Merci 18qw4z Invasiveness
  • Proved invasive outside its native range
  • Has a broad native range
  • Abundant in its native range
  • Highly adaptable to different environments
  • Is a habitat generalist
  • Tolerates, or benefits from, cultivation, browsing pressure, mutilation, fire etc
  • Tolerant of shade
  • Capable of securing and ingesting a wide range of food
  • Highly mobile locally
  • Benefits from human association (i.e. it is a human commensal)
  • Long lived
  • Fast growing
  • Has high reproductive potential
  • Reproduces asexually
Impact outcomes
  • Loss of medicinal resources
  • Monoculture formation
  • Reduced native biodiversity
  • Threat to/ loss of native species
Impact mechanisms
  • Causes allergic responses
  • Competition - monopolizing resources
  • Competition - smothering
  • Competition
  • Pest and disease transmission
  • Rapid growth
  • Rooting
Likelihood of entry/control
  • Highly likely to be transported internationally deliberately

Uses

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C. esculenta, commonly known in its crop form as taro, is mainly cultivated for its starchy stem tubers and corms. These corms are eaten boiled, fried or roasted as a side dish or are used to produce starch and flour. Taro corm puree makes an easily digested, low-allergenic baby food. In times of scarcity, this species is used as a famine food and in some regions leaves are used for food after cooking.

Waste leaves, corms and peel can be cooked or fermented into silage for animal feed. Most taro in South-East Asia is consumed by humans, but it also has uses in religious festivals and in folk medicines and is fed to livestock, primarily pigs.

In areas of Indonesia where rice is not grown, taro is eaten as a staple, baked, boiled or cooked in bamboo tubes. In Java, confections are prepared from taro flavoured with coconut and sugar: fried taro slices and taro chips are popular snacks. The leaves are used in preparing 'buntil' (salted fish with spices, grated coconut and vegetables, wrapped up and steamed in a taro leaf), and petioles are cooked. In Malaysia, taro is cooked in similar ways and also plays a role in religious festivals. Leaves are boiled and eaten as salad with spicy sauce, and petioles are cooked with coconut milk, meat and prawns. Taro in the Philippines is used primarily when more popular starches and green vegetables are in short supply. Corms are boiled, chipped and fried or made into confections. In Hawaii and parts of Polynesia, the corms are cooked and pounded into a paste that is allowed to ferment to produce 'poi'. A steamed pudding is made from grated taro and coconut.

Fibre obtained from the leaf stalk has been used for plaiting. In Asia and Africa, this species is also used in traditional medicine to treat arterial hypertension, liver problems, ulcers, snakebites, and rheumatism (Onwueme, 1999; Safo-Kantaka, 2004).

Uses List

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Animal feed, fodder, forage

  • Fodder/animal feed

Environmental

  • Amenity

General

  • Ornamental
  • Sociocultural value

Human food and beverage

  • Beverage base
  • Flour/starch
  • Food additive
  • Root crop
  • Vegetable

Materials

  • Fibre

Medicinal, pharmaceutical

  • Traditional/folklore

Similarities to Other Species/Conditions

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C. esculenta may be confused with other plants having large arrowhead-shaped leaf blades, such as species in the genera Alocasia and Xanthosoma (i.e., Xanthosoma sagittifolium). However, leaves of all similar-looking species are not peltate (i.e., their petioles are attached at the leaf-blade margin, rather than in the middle as with C. esculenta; Langeland et al., 2008).

Prevention and Control

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Physical/Mechanical Control

Mechanical removal of C. esculenta may be effective but is labour intensive. All corms and tubers should be removed to prevent spread. Dense stands should be removed using specialized machinery.

Chemical Control

Chemical control of C. esculenta should include mechanical cutting of the plants to the base followed by applications of N-phosphonomethyl-glycine (glyphosate). Follow-up treatments are recommended at 6 weeks intervals until control is completed (Queensland Department of Primary Industries and Fisheries, 2011). This procedure is not recommended when plants are growing near rivers or streams.

Bibliography

White T Heart Whistles Print shirt Merci 18qw4z Ghani FD, 1980. The status of keladi China, Colocasia esculenta (L.) Schott, cultivation in Peninsula Malaysia. IFS Provisional Report No 5. Stockholm, Sweden: International Foundation for Science, 35-54.

Ghani FD, 1981. Conservation and utilization of Colocasia (cultivars) and edible aroids (keladi) in Malaysia. Tropical Root and Tuber Crops Newsletter, 12, 13:38-46.

Sastrapradja S, Hambali GG, 1980. Taro (Colocasia spp.) as a source of carbohydrate, vitamins and minerals in Indonesian diets. IFS Provisional Report No 5. Stockholm, Sweden: International Foundation for Science, 17-28.

Villanueva MR, Tupas GL, 1980. Taro production in the Philippines: its prospects and problems. IFS Provisional Report No 5. Stockholm, Sweden: International Foundation for Science, 99-111.

Wang J-K, ed, 1983. Taro, a review of Colocasia esculenta and its potentials. Hawaii, USA: University of Hawaii Press.

Wilson JE, 1984. Cocoyam. In: Goldsworthy PR, Fisher NM, eds. The physiology of tropical field crops. Chichester, UK: John Wiley and Sons, 589-605.

References

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Acebes JR, Arco Mdel, García-Gallo A, León MC, Pérez PL, Rodríguez O, Wildpret Torre Wde la, Martín VE, Marrero MC, Rodríguez ML, 2001. Pteridophyta and Spermatophyta. (Pteridophyta y Spermatophyta.) In: Lista de especies silvestres de Canarias (hongos, plantas y animales terrestres) [ed. by Izquierdo, I. \Martín, J. L. \Zurita, N. \Arechavaleta, M.]., Spain: Consejería de Medio Ambiente y Ordenación Territorial, Gobierno de Canarias, 96-143

Acevedo-Rodríguez P, Strong MT, 2005. Monocots and Gymnosperms of Puerto Rico and the Virgin Islands. Contributions from the United States National Herbarium, volume 52:415 pp

Acevedo-Rodríguez P, Strong MT, 2012. Catalogue of the Seed Plants of the West Indies. Smithsonian Contributions to Botany, 98:1192 pp. Washington DC, USA: Smithsonian Institution. http://botany.si.edu/Antilles/WestIndies/catalog.htm

Bello D, 1883. [English title not available]. (Apuntes para la flora de Puerto Rico. Segunda parte. Monoclamídeas.) Anales de la Sociedad Española de Historia Natural, 12:103-130

Brako L, Zarucchi JL, 1993. Catalogue of the flowering plants and gymnosperms of Peru (Monographs in Systematic Botany from the Missouri Botanical Garden, vol. 45), 1-1286

Broome R, Sabir K, Carrington S, 2007. Plants of the Eastern Caribbean. Online database. Barbados: University of the West Indies. http://ecflora.cavehill.uwi.edu/index.html

Carney JA, Rosomoff RN, 2009. In the Shadow of Slavery: Africa's Botanical Legacy in the Atlantic World. Berkeley, California, USA: University of California Press, 280 pp

Chacón E, Saborío G, 2012. Red Interamericana de Información de Especies Invasoras, Costa Rica ([English title not available]). San José, Costa Rica: Asociación para la Conservación y el Estudio de la Biodiversidad. http://invasoras.acebio.org

Charles Darwin Foundation, 2008. Database inventory of introduced plant species in the rural and urban zones of Galapagos. Galapagos, Ecuador: Charles Darwin Foundation

Cho JJ, Yamakawa RA, Hollyer J, 2007. Hawaiian kalo, past and future. Sustainable Agriculture, SA-1. Hawaii, USA: College of Tropical Agriculture and Human Resources. https://www.ctahr.hawaii.edu/oc/freepubs/pdf/SA-1.pdf

Chong KY, Tan HTW, Corlett RT, 2009. A checklist of the total vascular plant flora of Singapore: native, naturalised and cultivated species. Singapore: Raffles Museum of Biodiversity Research, National University of Singapore, 273 pp. http://lkcnhm.nus.edu.sg/nus/pdf/PUBLICATION/LKCNH%20Museum%20Books/LKCNHM%20Books/flora_of_singapore_tc.pdf

Coates DJ, Yen DE, Gaffey PM, 1988. Chromosome variation in taro, Colocasia esculenta: implications for origin in the Pacific. Cytologia, 53(3):551-560

Correa A, Galdames MDC, Stapf MNS, 2004. Catalogue of vascular plants of Panama (Catalogo de Plantas Vasculares de Panama.), Panama: Smithsonian Tropical Research Institute, 599 pp

DAISIE, 2012. Delivering Alien Invasive Species Inventories for Europe. DAISIE (online). www.europe-aliens.org

Ellison JC, 1990. Vegetation and floristics of the Tongatapu outliers. Atoll Research Bulletin, 332:36 pp. http://www.sil.si.edu/digitalcollections/atollresearchbulletin/issues/00332.pdf

Elzebroek, T., Wind, K., 2008. Guide to cultivated plants, CABI.vii-xi + 516 pp.

EPPO, 2014. PQR database. Paris, France: European and Mediterranean Plant Protection Organization. http://www.eppo.int/DATABASES/pqr/pqr.htm

Flora of China Editorial Committee, 2012. Flora of China Web. Cambridge, USA: Harvard University Herbaria. http://flora.huh.harvard.edu/china/

Florence J, Chevillotte H, Ollier C, Meyer JY, 2011. [English title not available]. (Base de données botaniques Nadeaud de l'Herbier de la Polynésie Française (PAP).) . http://www.herbier-tahiti.pf

Florida Exotic Pest Plant Council, 2011. Florida EPPC's 2011 Invasive Plant Species List. http://www.fleppc.org/list/11list.html

Fosberg FR, Sachet M-H, Oliver RL, 1987. A geographical checklist of the Micronesian monocotyledonae. Micronesia, 20:1026

Funk V, Hollowell T, Berry P, Kelloff C, Alexander SN, 2007. Checklist of the plants of the Guiana Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contributions from the United States National Herbarium, 584 pp

González-Torres LR, Rankin R, Palmarola A (eds), 2012. Invasive plants of Cuba. (Plantas Invasoras en Cuba.) Bissea: Boletin sobre Conservacion de Plantad del Jardin Botanico Nacional, 6:1-140

Govaerts R, 2012. World Checklist of Araceae. London, UK: Royal Botanic Gardens, Kew. http://apps.kew.org/wcsp/

Graveson R, 2012. Plants of Saint Lucia. http://www.saintlucianplants.com

Greenwell AB, 1947. Taro - with special reference to its culture and uses in Hawaii. Economic Botany, 1:276-89

Grisebach AHR, 1864. Flora of the British West Indies Islands. London, UK: Lovell Reeve & Co., 789 pp

Groves RH, Boden R, Lonsdale, WM, 2005. Jumping the Garden Fence: Invasive Garden Plants in Australia and their environmental and agricultural impacts. Sydney, Australia: WWF. http://www.wwf.org.au/news_resources/resource_library/?1928/Jumping-the-Garden-Fence-Invasive-garden-plants-in-Australia-and-their-environmental-and-agricultural-impacts

Hancock IR, Henderson CP, 1988. Flora of the Solomon Islands. Research Bulletin No. 7. Honiara, Solomon Islands: Dodo Creek Research Station

Hokche O, Berry PE, Huber O, 2008. Nuevo Catálogo de la Flora Vascular de Venezuela (New catalogue of the vascular flora of Venezuela). Caracas, Venezuela: Fundación Instituto Botánico de Venezuela, 860 pp

Holm L, Pancho JV, Herberger JP, Plucknett DL, 1979. A Geographical Atlas of World Weeds. Toronto, Canada: John Wiley and Sons Inc

Idárraga-Piedrahita A, Ortiz RDC, Callejas Posada R, Merello M, 2011. Flora of Antioquia. (Flora de Antioquia.) Catálogo de las Plantas Vasculares, vol. 2. Listado de las Plantas Vasculares del Departamento de Antioquia:939 pp

Jørgensen PM, León-Yànez S, 1999. Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard, 75. i-viii, 1-1182

Langeland KA, Cherry HM, McCormick CM, Craddock Burks KA, 2008. Identification and Biology of Non-native Plants in Florida's Natural Areas. Gainesville, Florida, USA: University of Florida IFAS Extension

Lebot, V., 2008. Tropical root and tuber crops: cassava, sweet potato, yams and aroids, CABI.xv-xix + 404 pp. http://www.cabi.org/cabebooks/ebook/20093020206 doi:10.1079/9781845934248.0000

Lorence DH, Flynn T, 2010. Checklist of the plants of Kosrae. Unpublished checklist. National Tropical Botanical Garden. Lawai, Hawaii: National Tropical Botanical Garden, 26

Loy TH, Spriggs M, Wickler S, 1992. Direct evidence for human use of plants 28,000 years ago: Starch residues on stone artefacts from the northern Solomon Islands. Antiquity, 66, 898-912.

Madagascar Catalogue, 2012. Catalogue of the Vascular Plants of Madagascar. Antananarivo, Madagascar: Missouri Botanical Garden, Madagascar Research and Conservation Program. http://www.efloras.org/madagascar

McCormack G, 2007. Cook Islands Biodiversity Database, Version 2007.2. Rarotonga: Cook Islands Natural Heritage Trust. http://cookislands.bishopmuseum.org

Meyer JY, 2007. Rapport de mission sur l'Ile d'Uvea (Wallis & Futuna) du 6 au 17 Novembre 2007: Inventaire preliminaire de la flore vasculaire secondaire ([English title not available]). Papeete, Tahiti: Ministère de l'Education, l'Enseignement Supérieur et la Recherche, 39 pp. http://www.li-an.fr/jyves/Meyer_2007_Rapport_Plantes_Introduites_Wallis.pdf

Molina RA, 1975. Enumeration of the plants of Honduras. (Enumeración de las plantas de Honduras) Ceiba, 19(1):1-118

Onwueme I, 1999. Taro cultivation in Asia and the Pacific. Bangkok, Thailand: Food and Agriculture Organization of the United Nations. Regional Office for Asia and the Pacific, 15 pp. [FAO Report.]

PIER, 2012. Pacific Islands Ecosystems at Risk. Honolulu, USA: HEAR, University of Hawaii. http://www.hear.org/pier/index.html

Proctor GR, 1984. Flora of the Cayman Islands. London, UK: HMSO, 834 pp

Queensland Department of Primary Industries and Fisheries, 2011. Special edition of Environmental Weeds of Australia for Biosecurity Queensland., Australia: The University of Queensland and Department of Primary Industries and Fisheries. http://keyserver.lucidcentral.org/weeds/data/03030800-0b07-490a-8d04-0605030c0f01/media/Html/Index.htm

Ragone D, Lorence DH, 2003. Botanical and Ethnobotanical Inventories of the National Park of American Samoa. Hawaii, USA: Pacific Cooperative Studies Unit, Department of Botany, University of Hawaii, 91 pp

Randall RP, 2012. A Global Compendium of Weeds. Perth, Australia: Department of Agriculture and Food Western Australia, 1124 pp. http://www.cabi.org/isc/FullTextPDF/2013/20133109119.pdf

Robin GC, 2008. Commercial dasheen (Colocasia esculenta (L.) Schott var. esculenta) production and post-harvest protocol for OECS., Dominica: Caribbean Agriculture Research and Development Institute, 30 pp

Safo-Kantaka O, 2004. Colocasia esculenta (L.) Schott. Record from Protabase. PROTA (Plant Resources of Tropical Africa / Ressources végétales de l'Afrique tropicale) [ed. by Grubben, G. J. H. \Denton, O. A.]. Wageningen, Netherlands: PROTA. http://database.prota.org/search.htm

Silva VRMda, 2002. [English title not available]. (Flora da Madeira Plantas Vasculares Naturalizadas no Arquipélago da Madeira.) Boletim do Museu Municipal do Funchal (História Natural), supplement, 8:5-281

Smith AC, 1979. Flora vitiensis nova. A new flora of Fiji. Hawaii, Pacific Tropical Botanical Garden, 1:495

Sreekumari MT, Mathew PM, 1991. Karyotypically distinct morphotypes in taro (Colocasia esculenta (L.) Schott). Cytologia, 56(3):399-402

Standley PC, Steyermark JA, 1958. Araceae. Fieldiana-Botany:304-363. [Flora of Guatemala.]

Stevens PF, 2012. Angiosperm Phylogeny Website. http://www.mobot.org/MOBOT/research/APweb/

Stone BC, 1970. The Flora of Guam. Micronesica, 6:1-659

Sykes WR, 1970. Contributions to the flora of Niue. New Zealand Department of Scientific and Industrial Research Bulletin 200. p. 238

Sykes WR, 1970. Contributions to the flora of Niue. New Zealand Department. Sci. Indust. Res. Bull. 200: 1-321

US Fish and Wildlife Service, 1995. San Marcos and Comal and Associated Aquatic Ecosystems (Revised) Recovery Plan. In: San Marcos and Comal and Associated Aquatic Ecosystems (Revised) Recovery Plan : US Fish and Wildlife Service.x + 93 pp. + 28 pp. of appendices.

USDA-ARS, 2012. Germplasm Resources Information Network (GRIN). Online Database. Beltsville, Maryland, USA: National Germplasm Resources Laboratory. https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomysearch.aspx

USDA-NRCS, 2012. The PLANTS Database. Baton Rouge, USA: National Plant Data Center. http://plants.usda.gov/

Villaseñor JL, Espinosa-Garcia FJ, 2004. The alien flowering plants of Mexico. Diversity and Distributions, 10(2):113-123

Wagner WL, Herbst DR, Sohmer SH, 1999. Manual of the flowering plants of Hawaii. Revised edition. Honolulu, Hawai'i, USA: Bishop Museum Press, 1919 pp

Whitney LD, Bowers FAI, Takahashi M, 1939. Taro varieties in Hawaii. Honolulu (HI): Hawaii Agricultural Experiment Station. 86 pp. (Bulletin No. 84)

Wu T, 2001. Check List of Hong Kong Plants. Hong Kong Herbarium and the South China Institute of Botany. Check List of Hong Kong Plants:384 pp. http://www.hkflora.com/v2/flora/plant_check_list.php

Zhengyi W, Raven PH, Hong D, 2011. Flora of China. Flora of China. http://flora.huh.harvard.edu/china/mss/intindex.htm

Links to Websites

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Website URL Comment
Angiosperm Phylogeny Website http://www.mobot.org/mobot/research/apweb/
FAO: Technical Report on Taro Cultivation in Asia and the Pacific http://www.fao.org/docrep/005/AC450E/ac450e00.htm#Contents
Flora of the West Indies http://botany.si.edu/antilles/WestIndies/
Florida Exotic Pest Plant Council http://www.fleppc.org/ID_book/Colocasia%20esculenta.pdf

Contributors

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22/02/13 Updated by:

Julissa Rojas-Sandoval, Department of Botany-Smithsonian NMNH, Washington DC, USA

Pedro Acevedo-Rodríguez, Department of Botany-Smithsonian NMNH, Washington DC, USA

Distribution Maps

White T Heart Whistles Print shirt Merci 18qw4z
You can pan and zoom the map

Material & care

Outer fabric material: 100% silk

Care instructions: Dry clean only

Details

Pattern: Plain

Article number: HU722C01G-K11

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